Meet our grantees
Wild Animal Initiative funds academic research on high-priority questions in wild animal welfare.
The goal of our grants program is to fund research that deepens scientific knowledge of the welfare of wild animals in order to better understand how to improve the welfare of as many wild animals as possible, regardless of what causes the threats to their well-being.
We showcase our grantees and their projects here and continuously update this page as new projects are added.
Validating behavioral, acoustic, and physiological indicators of welfare in urban birds
Grantee: Miriam Soledad Vazquez
Institution: Universidad Nacional del Sur, Argentina
Project summary
This project will evaluate whether behavioral, acoustic, and physiological indicators consistently reflect welfare-relevant experiences in two urban thrush species. Welfare will be assessed by combining focal behavioral observations (vigilance, aggression, displacement, foraging activity, and tolerance of conspecifics and heterospecifics), passive acoustic monitoring (focal activity rates and calling patterns), and feather glucocorticoid analyses across urban sites that differ in vegetation cover and human disturbance. These indicators will be interpreted together to evaluate whether they show concordant, biologically meaningful responses across urban environments, while recognizing that different indicators may capture different temporal or contextual dimensions of affective state and welfare.
Grantee: Miriam Soledad Vazquez
Institution: Universidad Nacional del Sur, Argentina
Grant amount: $9,600
Grant type: Seed grants
Focal species: Patagonian thrush (Turdus falcklandii), Rufous-bellied thrush (T. rufiventris)
Conservation status: N/A: Multiple focal species
Disciplines: Ornithology, animal behavior, physiology
Research location: Argentina
Project summary
This project will evaluate whether behavioral, acoustic, and physiological indicators consistently reflect welfare-relevant experiences in two urban thrush species that have recently expanded their ranges into Argentine cities. Welfare will be assessed by combining focal behavioral observations (vigilance, aggression, displacement, foraging activity, and tolerance of conspecifics and heterospecifics), passive acoustic monitoring (focal activity rates and calling patterns), and feather glucocorticoid analyses across urban sites that differ in vegetation cover and human disturbance. These indicators will be interpreted together to evaluate whether they show concordant, biologically meaningful responses across urban environments, while recognizing that different indicators may capture different temporal or contextual dimensions of affective state and welfare.
Why we funded this project
The study provides a proof of concept for integrating behavioral observations, passive acoustic monitoring, and physiological measures within a multidimensional welfare assessment framework applied under natural conditions. It also addresses whether commonly proposed welfare indicators remain informative across different species and environmental contexts, including urban environments shaped by human disturbance and novel biotic interactions.
Evaluating ice nests: Effects of a welfare intervention on long-lived insect societies and their guests
Grantee: Christina Kwapich
Institution: University of Central Florida, United States
Project summary
When social organisms are translocated, the destruction of multi-generational nests, loss of familiar territory, and disruption of demography could negatively influence the welfare of society members, future generations, and dependent symbionts. This project will translocate Florida harvester ant (Pogonomyrmex badius) colonies by burying replica nests made from ice. The team will measure risk-averse behaviors in ants across generations, as well as the stability of colony foraging routes, demography, and the re-colonization of symbiont populations. By monitoring ants in freshly transplanted and historically transplanted colonies, the project will provide data on the effects of social disruption on ant societies.
Grantee: Christina Kwapich
Institution: University of Central Florida, United States
Grant amount: $10,000
Grant type: Seed grants
Focal species: Florida harvester ant (Pogonomyrmex badius) and others
Conservation status: Not evaluated
Disciplines: Entomology, wildlife management
Research location: United States
Project summary
US laws increasingly require that species of interest be removed from land marked for development and translocated to suitable sites. For social organisms, the destruction of multi-generational nests, loss of familiar territory, and disruption of demography could negatively influence the welfare of surviving society members, future generations, and dependent symbionts. This project will translocate a population of Florida harvester ant (Pogonomyrmex badius) colonies by burying replica nests made from ice. The team will measure risk-averse behaviors in ants across generations, as well as the stability of colony foraging routes, demography, and the re-colonization of symbiont populations. By monitoring ants in freshly transplanted and historically transplanted colonies, the project will provide data on the effects of social disruption on long-lived animal societies made up of short-lived members.
Why we funded this project
The “ice nest” technique represents a welfare intervention that could mitigate some of the negative effects of social disruption associated with research or land development. By publishing on the feasibility of the ice nest technique, social insect well-being may be given serious consideration in future environmental mitigation and research protocols.
Assessing behavioral and physiological welfare indicators for the Kashmir loach (Triplophysa kashmirensis)
Grantee: Misba Rehman
Institution: Sher-e-Kashmir University of Agricultural Sciences and Technology, India
Project summary
This project aims to develop and validate non-invasive behavioral and physiological welfare indicators for the Kashmir loach (Triplophysa kashmirensis). The researchers will use underwater cameras to record behaviors like activity, refuge use, foraging, and abnormal swimming patterns. Physiological welfare will be assessed using mucus sampling to measure cortisol concentrations and supplementary indicators including oxidative stress and immune markers. The team will validate indicators by testing their correlation with key water quality parameters. The project aims to establish the first field-based welfare assessment framework for this species and develop a transferable model for wild freshwater fishes inhabiting mountain stream ecosystems.
Grantee: Misba Rehman
Institution: Sher-e-Kashmir University of Agricultural Sciences and Technology, India
Grant amount: $6,500
Grant type: Seed grants
Focal species: Kashmir Loach (Triplophysa kashmirensis)
Conservation status: Critically endangered
Disciplines: Ichthyology, physiology, animal behavior
Research location: India
Project summary
This project aims to develop and validate non-invasive behavioral and physiological welfare indicators for the Kashmir loach (Triplophysa kashmirensis), a small benthic freshwater fish endemic to Himalayan streams. Fish will be observed under natural stream conditions, with underwater cameras recording behaviors like activity, refuge use, foraging, and abnormal swimming patterns. Physiological welfare will be assessed using mucus sampling to measure cortisol concentrations and supplementary indicators including oxidative stress and immune markers. Indicators will be validated by testing their correlation with key water quality parameters. By integrating behavioral, physiological, and environmental data, the project aims to establish the first field-based welfare assessment framework for this species and develop a transferable model for wild freshwater fishes inhabiting mountain stream ecosystems.
Why we funded this project
This project contributes to wild animal welfare science by developing and validating a field-applicable, non-invasive framework for assessing welfare in free-living freshwater fishes, a group that remains highly understudied in welfare research.
Decoding aquatic mollusk welfare: an investigation of potential physiological and cognitive indicators
Grantee: Laura Webb
Institution: Wageningen University, Netherlnads
Project summary
This project aims to explore physiological and cognitive indicators of welfare in two species of aquatic mollusk: highly mobile cuttlefish (dwarf cuttlefish; Ascarosepion bandense) and sessile (blue) mussels (Mytilus edulis). Valid indicators of mollusk welfare can help us better understand their welfare in changing environmental conditions, such as those associated with climate change. The researchers will explore heart rate variability and respiratory variability as possible physiological indicators, and optimism in a judgment bias test as a cognitive indicator of welfare. Following this, successful metrics can in turn be used to explore the impact of environmental changes on these species' welfare.
Grantee: Laura Webb
Institution: Wageningen University, Netherlands
Grant amount: $10,000
Grant type: Seed grants
Focal species: Bobtail squid (Euprymna berryi), dwarf cuttlefish (Ascarosepion bandense), blue mussel (Mytilus edulis)
Conservation status: Not evaluated
Disciplines: Malacology, cognition, physiology
Research location: Netherlands
Project summary
This project aims to explore physiological and cognitive indicators of welfare in two species of aquatic mollusk: highly mobile cuttlefish (dwarf cuttlefish; Ascarosepion bandense) and sessile (blue) mussels (Mytilus edulis). Valid indicators of mollusk welfare can help us better understand their welfare in changing environmental conditions, such as those associated with climate change. The researchers will explore heart rate variability (mussels) and respiratory variability as possible physiological indicators, and optimism in a judgment bias test as a cognitive indicator of welfare (cuttlefish). Following this, successful metrics can in turn be used to explore the impact of environmental changes on these species' welfare.
Why we funded this project
This project is highly innovative and exploratory in nature, focused on neglected species in welfare research, with sentience accepted for cuttlefish in EU law but not yet for mussels. This work will raise attention to the topic of aquatic mollusk welfare and possible assessment methods in these species.
Swan Lake: Assessing the welfare of urban mute swans
Grantee: Irena Uzelac
Institution: Animal Rescue Serbia
Project summary
This project investigates how human-provided food, environmental conditions, and injuries affect the welfare of mute swans in urban Belgrade. The project will assess welfare using indicators such as body condition, visible deformities, injury rates, and behavior patterns like foraging and aggression. By comparing sites with different levels of human feeding and variations in temperature, the project aims to understand how these factors shape swan health and behavior. Reproductive success and juvenile survival will also be monitored to evaluate long-term welfare outcomes. A mix of behavioral observation, environmental monitoring, and welfare scoring will be used, integrating both natural and anthropogenic stressors. The ultimate goal is to develop ecologically valid welfare indicators that reflect real-world challenges faced by urban wildlife.
Grantee: Irena Uzelac
Institution: Animal Rescue Serbia, Serbia
Grant amount: $10,000
Grant type: Seed grants
Focal species: Mute swan (Cygnus olor)
Conservation status: Least concern
Disciplines: Ornithology, animal behavior, physiology
Research locations: Serbia
Project summary
This project investigates how human-provided food, environmental conditions, and injuries affect the welfare of mute swans (Cygnus olor) in urban Belgrade. The project will assess welfare using indicators such as body condition, visible deformities (e.g., angel wing), injury rates, and behavior patterns like foraging and aggression. By comparing sites with different levels of human feeding and variations in temperature, the project aims to understand how these factors shape swan health and behavior. Reproductive success and juvenile survival will also be monitored to evaluate long-term welfare outcomes. A mix of behavioral observation, environmental monitoring, and welfare scoring will be used, integrating both natural and anthropogenic stressors. The ultimate goal is to develop ecologically valid welfare indicators that reflect real-world challenges faced by urban wildlife.
Why we funded this project
This project will help address the question of how food from humans and urban environmental stressors affect the health and welfare of wild birds by applying practical welfare indicators in a free-living, urban population of mute swans. While this question has been longstanding, most research has been limited to narrower health (disease) or demographic impacts. We hope that this project will provide information on when and how anthropogenic food might be a welfare threat or opportunity.
Validating body posture as a novel marker of well-being in animals
Grantee: Nicola Koyama
Institution: Liverpool John Moores University
Project summary
A few studies on nonhuman primates have found that a hunched posture is a response to social separation and physical inflammation, but research linking whole body posture and physiological measures of welfare is lacking. This project will combine established measures of affective valence (nasal temperature) and arousal (iris-pupil ratio) to validate body posture as a new measure of affective valence. Two months of video data will be collected and used for thermal imaging, behavioral, and postural analysis from groups of wild Barbary macaques, along with pilot data for comparison from non-human primate zoo animals. Ultimately, the project aims to share a new validated measure and conceptual framework that can be applied to a range of wild mammals.
Grantee: Nicola Koyama
Institution: Liverpool John Moores University, United Kingdom
Grant amount: $9,995
Grant type: Seed grants
Focal species: Barbary macaque (Macaca sylvanus)
Conservation status: Endangered
Disciplines: Primatology, animal behavior, physiology
Research locations: United Kingdom, Morocco
Project summary
A few studies on non-human primates have found that a hunched posture is a response to social separation and physical inflammation, but research linking whole body posture and physiological measures of welfare is lacking. This project will combine established measures of affective valence (nasal temperature) and arousal (iris-pupil ratio) to validate body posture as a new measure of affective valence. Two months of video data will be collected and used for thermal imaging, behavioral, and postural analysis from groups of wild Barbary macaques, along with pilot data for comparison from non-human primate zoo animals. Ultimately, the project aims to share a new validated measure and conceptual framework that can be applied to a range of wild mammals.
Why we funded this project
This project will validate an indicator for assessing welfare that could be relevant across primates, and provide proof of concept for its application in the field.
An Evaluation of Effective Disturbance Mitigation Measures for European Badgers (Meles meles) Occupying Setts During Forestry Operations
Grantee: Stephen McAuliffe
Institution: University of Brighton and Forest Research
Project summary
In order to avoid possible disturbance to European badgers, forestry guidance in England states that activities using heavy machinery should be prohibited within 20 meters of active badger setts. This research aims to validate that the approved noise and vibration mitigation buffer distances intended to prevent disturbance from anthropogenic activities are effective. Observations of badger behavior will be made in woodland habitats, and fecal cortisol metabolites will be collected for analysis. If changes in behavior and increases in cortisol levels are not found following exposure to forestry noise and vibration greater than 20 meters away, robust evidence will be provided to validate that badger welfare is not being adversely impacted by legally sanctioned forestry activities.
Grantee: Stephen McAuliffe
Institution: University of Brighton and Forest Research, United Kingdom
Grant amount: $10,137
Grant type: Seed grants
Focal species: European badger (Meles meles)
Conservation status: Least concern
Disciplines: Animal behavior, physiology, mammalogy, human-wildlife conflicet
Research locations: United Kingdom
Project summary
In order to avoid possible disturbance to European badgers (Meles meles), forestry guidance in England states that activities using heavy machinery should be prohibited within 20 meters of active badger setts. However, this commonly adopted mitigation measure has never been scientifically assessed. As a result, it relies on the untested assumption that noise and vibrations do not disturb badgers if a buffer zone of 20 meters is maintained between forestry operations and setts. This research aims to validate that the approved noise and vibration mitigation buffer distances intended to prevent disturbance from anthropogenic activities are effective. Observations of badger behavior will be made in woodland habitats, and fecal cortisol metabolites will be collected for analysis. If changes in behavior and increases in cortisol levels are not found following exposure to forestry noise and vibration greater than 20 meters away, robust evidence will be provided to validate that badger welfare is not being adversely impacted by legally sanctioned forestry activities.
Why we funded this project
This project will help us understand the welfare impacts of anthropogenic disturbance on a common mammal, and could lead to a very near-term intervention to mitigate them if the project results in a recommendation to increase buffer distances. We hope it will set a precedent for testing and improving animal welfare protection methods routinely advised and adopted by many land-based sectors such as forestry, utilities providers, construction and agriculture.
Assessment of welfare impacts of parasites and pathogens on migratory flamingos among Salt Lakes of East Africa
Grantee: Robert Modest Byamungu
Institution: Sokoine University of Agriculture
Project summary
This project will investigate the effects of parasites and pathogens on the welfare of two flamingo species ― the greater flamingo and the lesser flamingo — in the Salt Lakes of East Africa using non-invasive methods. To achieve this, the project will establish two sites: an experimental site and a control site. At both sites, it will (i) estimate parasites and pathogens loads, (ii) assess stress behaviour, (iii) quantify stress physiology and, (iv) monitor food resources of flamingos. Welfare will be assessed through behaviors (e.g., body scratching and feathers plucking), as well as signs of stress (e.g., fecal cortisol levels). The project helps provide an understanding on how environmental pressures impact wild animals' quality of life.
Grantee: Robert Modest Byamungu
Institution: Sokoine University of Agriculture, Tanzania
Grant amount: $9,920
Grant type: Seed grants
Focal species: Greater flamingo (Phoenicopterus roseus)
Conservation status: Least concern
Disciplines: Ornithology, animal behavior, physiology, infectious disease
Research locations: Tanzania
Project summary
This project seeks to deepen understanding of flamingo migration among the Salt Lakes of East Africa by looking at the role of parasites and pathogens on the welfare of two flamingo species ― the greater flamingo and the lesser flamingo — using non-invasive methods. To achieve this, the project will establish two sites: an experimental site and a control site. At both sites, it will (i) estimate parasites and pathogens loads, (ii) assess stress behaviour, (iii) quantify stress physiology and, (iv) monitor food resources of flamingos. Welfare will be assessed through behaviors (e.g., body scratching and feathers plucking), as well as signs of stress (e.g., fecal cortisol levels). The project helps provide an understanding on how environmental pressures impact wild animals' quality of life.
Why we funded this project
Flamingos are a highly numerous species, so the findings of this project have the potential to help researchers understand the welfare of a large number of individuals. Migration of other group-living animals may also be influenced by welfare, so the results may also be translatable to other species.
Validation of non-invasive measures for life stage in wild octopuses and AI-Assisted Behavioral Analysis
Grantee: Michaella Andrade
Institution: Federal University of ABC
Project summary
This project aims to develop non-invasive methods to assess the life stage and welfare of wild octopuses. The project will develop an AI approach that learns to recognize complex behaviors, such as body patterns and ventilation rates, in video footage. In addition, the project will develop a methodology to automatically measure the life stage of octopuses in videos and non-invasive in situ measurements of the distance between the octopuses’ eyes and eyeballs. The main goal is to correlate these metrics with different behavioral contexts to determine whether they can be used as reliable indicators of the animal’s affective state. In the long term, the project aims to create an automated tool that accelerates and improves behavioral data analysis.
Grantee: Michaella Andrade
Institution: Federal University of ABC, Brazil
Grant amount: $10,000
Grant type: Seed grants
Focal species: Octopuses (Octopus insularis)
Conservation status: Not evaluated
Disciplines: Animal behavior
Research locations: Brazil
Project summary
This project aims to develop non-invasive methods to assess the life stage and welfare of wild octopuses. The project will develop an innovative AI approach that automatically learns to recognize complex behaviors, such as body patterns and ventilation rates, directly from pre-recorded videos in the field. In addition, the project will develop a methodology to automatically measure the life stage of octopuses in pre-recorded videos based on our database and non-invasive in situ measurements of the distance between the octopuses’ eyes and eyeballs. The main goal is to correlate these metrics with different behavioral contexts to determine whether they can be used as reliable indicators of the animal’s affective state. In the long term, the project aims to create an automated tool that accelerates and improves behavioral data analysis, enabling large-scale studies of the welfare of wild octopuses.
Why we funded this project
This project extends a previous WAI-funded project and uses innovative approaches to analyzing behavioral data for welfare assessment.
Between freedom and security, what makes a feral pigeon happier?
Grantee: Brandon Mak
Institution: Technical University of Munich
Project summary
Feral pigeons in Germany are subject to the Augsburg model of non-lethal population control, which involves attracting them to breed in dovecots where food and nest sites are provided, and their eggs exchanged with dummies. Others are captured and housed in aviaries under similar conditions. This project investigates the welfare implications of Germany’s non-lethal population control strategies. This is done by comparing the stress profiles of feral pigeons living “wild” in the streets, free-living pigeons breeding and feeding in dovecots, and formerly free-living pigeons translocated into aviaries. Fieldwork will be conducted to weigh pigeons (body mass measurement), collect egg and feather samples from their nests (corticosterone concentrations), and observe their behavior (locomotion and aggression).
Grantee: Brandon Mak
Institution: Technical University of Munich, Germany
Grant amount: $10,900
Grant type: Seed grants
Focal species: Feral pigeon/rock dove (Columba livia domestica)
Conservation status: Least concern
Disciplines: Ornithology, animal behavior, physiology, population ecology, wildlife management
Research locations: Germany
Project summary
Feral pigeons in Germany are subject to the Augsburg model of non-lethal population control, which involves attracting them to breed in dovecots where food and nest sites are provided, and their eggs exchanged with dummies. Others are captured and housed in aviaries under similar conditions. While aimed at reducing pigeon populations, providing food, and reducing exposure to predators, it’s possible that these measures may incidentally alleviate chronic stress in free-living individuals. This project investigates the welfare implications of Germany’s non-lethal population control strategies. This is done by comparing the stress profiles of feral pigeons living “wild” in the streets, free-living pigeons breeding and feeding in dovecots, and formerly free-living pigeons translocated into aviaries. Fieldwork will be conducted to weigh pigeons (body mass measurement), collect egg and feather samples from their nests (corticosterone concentrations), and observe their behavior (locomotion and aggression).
Why we funded this project
This project will contribute to our understanding of the potential welfare effects of (non-chemical) fertility control in pigeons, a leading candidate for near-term interventions to improve welfare in urban contexts. It will also provide some information on the validity of egg corticosterone as a non-invasive indicator of stress by measuring it alongside other more established methods (feather corticosterone, morphological and behavioural indicators) for a more robust welfare assessment.