Meet our grantees
Wild Animal Initiative funds academic research on high-priority questions in wild animal welfare.
The goal of our grants program is to fund research that deepens scientific knowledge of the welfare of wild animals in order to better understand how to improve the welfare of as many wild animals as possible, regardless of what causes the threats to their well-being.
We showcase our grantees and their projects here and continuously update this page as new projects are added.
Acoustic indicators of welfare in neotropical frogs: Calls as early-warning signals of sublethal suffering related to infection and environmental stress
Grantee: Erin Wall
Institution: Wavelength Animal Communication Science and University of Texas at Austin, United States
Project summary
Emerging evidence indicates that frog call characteristics are modulated by environmental and physiological conditions, offering the potential for rapid insights into an individual’s welfare. This project will examine the relationship between environmental conditions, physiology, and behavior to connect indices of welfare to variation in acoustic signals in neotropical frogs in Panama. By linking these measures to natural levels of sublethal infection and variation in environmental conditions, the researchers will aim to identify acoustic signatures of welfare and develop a rapid, non-intrusive welfare detection tool for thousands of species of vocalizing frogs.
Grantee: Erin Wall
Institution: Wavelength Animal Communication Science and University of Texas at Austin, United States
Grant amount: $180,000
Grant type: Fellowship
Focal species: Túngara frog (Engystomops pustulosus), hourglass treefrog (Dendropsophus ebraccatus), leaf-litter toad (Rhinella alata), and others
Conservation status: Least concern
Disciplines: Herpetology, physiology, animal behavior
Research locations: Canada, Panama, United States
Project summary
Climate change, habitat loss, and infectious disease are creating compounding challenges for frogs, yet we have limited tools to monitor the behavioral and physiological consequences of those challenges, assess frog welfare, or detect sublethal suffering. Emerging evidence indicates that frog call characteristics are modulated by environmental and physiological conditions, offering the potential for rapid insights into an individual’s welfare. This project will examine the relationship between environmental conditions, physiology, and behavior to connect indices of welfare to variation in acoustic signals in neotropical frogs in Panama. By linking these measures to natural levels of sublethal infection and variation in environmental conditions, we aim to identify acoustic signatures of welfare and develop a rapid, non-intrusive welfare detection tool for thousands of species of vocalizing frogs.
Why we funded this project
Along with supporting an early-career researcher, this grant will deliver a behavioral assessment framework and acoustic classifier tool for anurans to deepen our understanding of welfare in this diverse taxon and address the need for evidence-based, scalable welfare assessment methods.
Developing, testing and validating indicators of wild insect welfare
Grantee: Carl Soulsbury
Institution: University of Lincoln, United Kingdom
Project summary
This project aims to develop and validate practical indicators of welfare in wild insects. Working with three common model species — Dubia cockroaches (Blaptica dubia), two-spotted crickets (Gryllus bimaculatus), and carrion beetles — it will measure a suite of behavioral and physiological stress responses including oxidative stress markers, fat reserves via NMR, and movement patterns tracked using AI-assisted video analysis. These measures will be validated against cognitive bias testing to identify which indicators reliably reflect negative welfare. The most field-accessible measures will be applied to wild carrion beetle populations, testing whether lab-derived welfare indicators hold up under natural conditions.
Grantee: Carl Soulsbury
Institution: University of Lincoln, United Kingdom
Grant amount: $92,148
Grant type: Challenge grants
Focal species: Carrion beetles (Nicrophorus vespilloides) and others
Conservation status: Least concern
Disciplines: Entomology, physiology, animal behavior, cognition
Research location: United Kingdom
Project summary
This project aims to develop and validate practical indicators of welfare in wild insects. Working with three common model species — Dubia cockroaches (Blaptica dubia), two-spotted crickets (Gryllus bimaculatus), and carrion beetles — it will measure a suite of behavioral and physiological stress responses including oxidative stress markers, fat reserves via NMR, and movement patterns tracked using AI-assisted video analysis. These measures will be validated against cognitive bias testing to identify which indicators reliably reflect negative welfare, and the most field-accessible measures will be applied to wild carrion beetle populations, testing whether lab-derived welfare indicators hold up under natural conditions. The goal is a generalizable welfare assessment toolkit that researchers can use across insect taxa in both captive and wild settings.
Why we funded this project
This project provides the first systematic, cross-taxa proof of concept for measuring welfare in wild insects, moving beyond the captive-only focus that has dominated invertebrate welfare research to date. Invertebrates make up the vast majority of animal life on Earth, yet almost nothing is known about their welfare in the wild, so the knowledge generated by this project has the potential to facilitate large-scale impact. Funding this project also deepens our work with an existing grantee.
Find Carl’s other project, studying red foxes and Eurasian hedgehogs, here.
Towards welfare-centered prescribed fire
Grantee: Dale Nimmo
Institution: Charles Sturt University, Australia
Project summary
This project will investigate how prescribed burning affects wild animal welfare, and whether welfare state helps determine survival and harm suffered. The team will study bush rats (Rattus fuscipes) in forest blocks scheduled for prescribed burns and in control sites, combining behavioral, body-condition, injury, and non-invasive physiological measures to infer welfare changes. Lightweight telemetry will enable tracking of survival and severe harm, while fire-severity mapping, refuge measurements, and camera traps will help identify the mechanisms involved. The goal is to generate evidence to help burns be planned and implemented in ways that reduce avoidable suffering.
Grantee: Dale Nimmo
Institution: Charles Sturt University, Australia
Grant amount: $98,700
Grant type: Challenge grants
Focal species: Bush rats (Rattus fuscipes) and others
Conservation status: N/A: Multiple focal species
Disciplines: Climate science, physiology, animal behavior, mammalogy
Research location: Australia
Project summary
This project will investigate how prescribed burning affects the welfare of wild animals, and whether an animal’s welfare state before exposure to fire helps determine survival and harm suffered. The research team will study bush rats (Rattus fuscipes) in forest blocks scheduled for prescribed burns and in control sites, sampling individuals before fire, shortly after fire, and during early recovery, combining behavioral, body-condition, injury, and non-invasive physiological measures to infer changes in affective state and welfare. Lightweight telemetry will enable tracking of survival and severe harm through the burn window and post-fire period, while fire-severity mapping, refuge measurements, and camera traps will help identify the mechanisms involved. The goal is to generate practical evidence for welfare-centered prescribed fire: burns planned and implemented in ways that retain refuges, maintain escape options, and reduce avoidable suffering.
Why we funded this project
By combining welfare indicators with telemetry, fire-exposure metrics, refuge measurements, and predator activity, the project helps move wild animal welfare beyond descriptive assessment toward identifying the mechanisms and management choices that could reduce suffering.
Environmental drivers of welfare in urban rats: A multimodal field study of micro-habitat conditions and affective behavior
Grantee: Emily Mackevicius
Institution: Basis Research Institute, United States
Project summary
This project will investigate how urban micro-habitat conditions shape the affective and social lives of brown rats (Rattus norvegicus). Better tools for understanding rat behavior could support both wild animal welfare science and integrative management practices. The project will first test whether indicators developed in laboratory rodents such as ultrasonic vocalizations, posture, movement, social behavior, and surface temperature can be used to infer affective state and social experience in urban field settings. The team will then model how environmental features predict these indicators, with the goal of identifying urban conditions associated with positive or negative experiences.
Grantee: Emily Mackevicius
Institution: Basis Research Institute, United States
Grant amount: $100,000
Grant type: Challenge grants
Focal species: Brown rat (Rattus norvegicus) and others
Conservation status: Least concern
Disciplines: Wildlife management, physiology, animal behavior
Research location: United States
Project summary
This project will investigate how urban micro-habitat conditions shape the affective and social lives of free-ranging brown rats (Rattus norvegicus). Rats are highly social, intelligent, and abundant animals, and better tools for understanding their behavior could support both wild animal welfare science and integrative management practices. The project will first test whether indicators developed in laboratory rodents such as ultrasonic vocalizations, posture, movement, social behavior, and surface temperature, can be used to infer affective state and social experience in urban field settings. The team will then model how environmental features predict these indicators, with the goal of identifying urban conditions associated with positive or negative experiences.
Why we funded this project
This project addresses a neglected but important question: how fine-scale features of human-built environments shape the subjective experiences of abundant urban animals. The resulting methods could support future welfare studies across sites, cities, and taxa, and provide tools for welfare-aware, evidence-based urban management.
Developing & refining behavioral assays for the measurement of ‘sentient welfare’ in gastropod molluscs
Grantee: Elizabeth Paul
Institution: University of Bristol, United Kingdom
Project summary
Compared to cephalopod mollusks, gastropod mollusks have been neglected in welfare research and public and legislative concern. This project will develop and refine key assays for the behavioral measurement of welfare in two gastropod species: the great pond snail (Lymnaea stagnalis) and the common garden snail (Cornu aspersum). The researchers will study the capacities of snails to experience consequences within two hypothesized domains of affective function: the first relating to a Punishment Avoidance System (associated with an animal’s responses to threats and punishments) and the second to a Reward Acquisition System (associated with an animal’s responses to opportunities and rewards).
Grantee: Elizabeth Paul
Institution: University of Bristol, United Kingdom
Grant amount: $49,476
Grant type: Discovery grants
Focal species: Great pond snail (Lymnaea stagnalis), common garden snail (Cornu aspersum)
Conservation status: Least concern
Disciplines: Sentience, animal behavior, malacology
Research location: United Kingdom
Project summary
The welfare of cephalopod mollusks is a topic of rapidly increasing scientific and public interest. But to date, their close relatives, the gastropod mollusks, have been almost completely neglected in welfare research, as well as in public and legislative concern. This project will develop and refine key assays for the behavioral measurement of welfare in two gastropod species: the great pond snail (Lymnaea stagnalis) and the common garden snail (Cornu aspersum). The researchers will focus on the capacities of snails to experience consequences within two hypothesized domains of affective function — the first relating to a Punishment Avoidance System (PAS; associated with an animal’s responses to threats and punishments) and the second to a Reward Acquisition System (RAS; associated with an animal’s responses to opportunities and rewards).
Why we funded this project
Extensive scientific attention has been paid to the pollutants and other anthropogenic challenges facing gastropod mollusks, but from the point of view of broader environmental damage and effects on vertebrate populations. This project extends that interest to the welfare of gastropod animals themselves. The research team hopes that this project will lead to a larger research program on molluscan welfare.
Feeling salty: A welfare framework for estuarine fishes
Grantee: Samantha Levell
Institution: New College of Florida, United States
Project summary
Estuaries are highly dynamic environments in which salinity can change rapidly. This project will study how short-term, ecologically realistic salinity shifts influence welfare in estuarine fishes using non-invasive measures such as waterborne cortisol, metabolic rate, exploration, shoaling behavior, and decision making. The project will focus on two common estuarine species: the sheepshead minnow (Cyprinodon variegatus) and the sailfin molly (Poecilia latipinna). By comparing juveniles and adults, the project aims to develop practical tools for assessing fish welfare in the wild and to better understand how environmental variability shapes the experiences of aquatic animals.
Grantee: Samantha Levell
Institution: New College of Florida, United States
Grant amount: $26,200
Grant type: Discovery grants
Focal species: Sheepshead minnow (Cyprinodon variegatus), sailfin molly (Poecilia latipinna), and others
Conservation status: N/A: Multiple focal species
Disciplines: Ichthyology, physiology, animal behavior
Research location: United States
Project summary
Estuaries are highly dynamic environments in which salinity can change rapidly. This project will study how short-term, ecologically realistic salinity shifts influence welfare in estuarine fishes using non-invasive measures such as waterborne cortisol, metabolic rate, exploration, shoaling behavior, and decision making. The project will focus on two common estuarine species: the sheepshead minnow (Cyprinodon variegatus) and the sailfin molly (Poecilia latipinna). By comparing juveniles and adults, the project aims to develop practical tools for assessing fish welfare in the wild and to better understand how environmental variability shapes the experiences of aquatic animals.
Why we funded this project
Most previous work on estuarine fishes has focused on survival and physiological tolerance rather than the animals’ lived experiences or affective states. By integrating non-invasive physiological and behavioral indicators, this project will help validate a practical framework for assessing fish welfare under ecologically realistic conditions.
Validating acoustic monitoring as a non-invasive welfare indicator in urban coyotes: Linking vocalizations to health status, social dynamics, and environmental quality
Grantee: Caroline Rowley
Institution: Duke University, United States
Project summary
Urban coyotes face welfare challenges including disease, parasites, habitat degradation, reliance on anthropogenic food sources, and stress from human-wildlife conflict. This project is a pilot study aimed at validating passive acoustic monitoring as a welfare assessment tool. Data will be collected via GPS collars fitted with microphones, autonomous recording units, thermal camera traps and biological sampling. Using models and machine learning, the researchers will test whether vocal features correlate with established welfare indicators and whether individual coyotes can be identified from their howls alone. The aim is to develop a scalable, non-invasive framework for monitoring the welfare of free-ranging animals.
Grantee: Caroline Rowley
Institution: Duke University, United States
Grant amount: $49,978
Grant type: Discovery grants
Focal species: Eastern coyote (Canis latrans)
Conservation status: Least concern
Disciplines: Animal behavior, mammalogy
Research location: United States
Project summary
Coyote populations have expanded dramatically across North America, but population-level success does not necessarily equate to good welfare. Urban coyotes in particular face welfare challenges including disease, parasites, habitat degradation, reliance on anthropogenic food sources, and stress from human-wildlife conflict. This project is a pilot study aimed at validating passive acoustic monitoring as a welfare assessment tool. Data will be collected via GPS collars fitted with microphones, autonomous recording units, thermal camera traps and biological sampling. Using models and machine learning, the researchers will test whether vocal features correlate with established welfare indicators and whether individual coyotes can be identified from their howls alone. The goal is to develop a scalable, non-invasive framework for monitoring the welfare of free-ranging animals.
Why we funded this project
By systematically linking acoustic features of coyote vocalizations to established welfare indicators across multiple domains, this study positions passive acoustic monitoring as a scalable, whole-animal welfare assessment tool that is likely transferable to other similar species using vocalizations for communication.
Comparative assessment of capybara (Hydrochoerus hydrochaeris) welfare in protected, semi-anthropized, and anthropized environments in Argentina
Grantee: Débora Silvia Racciatti
Institution: Universidad de Buenos Aires, Argentina
Project summary
This project will assess how different levels of human disturbance are associated with behavioral, physiological, and health-related welfare indicators in capybara (Hydrochoerus hydrochaeris). Using camera traps, direct observations, non-invasive fecal sampling, and body condition assessments, the team will evaluate a suite of indicators and test not only whether they vary in predicted directions across environments, but how consistently they converge across welfare domains, identifying which measures most reliably discriminate conditions of higher versus lower welfare in free-ranging populations.
Grantee: Débora Silvia Racciatti
Institution: Universidad de Buenos Aires, Argentina
Grant amount: $45,400
Grant type: Discovery grants
Focal species: Capybara (Hydrochoerus hydrochaeris)
Conservation status: Least concern
Disciplines: Human-wildlife conflict, physiology, animal behavior, mammalogy
Research location: Argentina
Project summary
This project compares capybara (Hydrochoerus hydrochaeris) populations living in protected, semi-anthropized, and highly anthropized environments in Argentina to assess how different levels of human disturbance are associated with behavioral, physiological, and health-related welfare indicators. Using camera traps, direct observations, non-invasive fecal sampling, and body condition assessments, the team will evaluate a suite of indicators and test not only whether they vary in predicted directions across environments, but how consistently they converge across welfare domains, identifying which measures most reliably discriminate conditions of higher versus lower welfare in free-ranging populations. By integrating validated measures into a comparative welfare assessment framework, the project aims to advance the empirical foundations of wild animal welfare science and generate evidence to inform welfare-aware wildlife management and coexistence strategies.
Why we funded this project
This project addresses a critical but underexplored question in wild animal welfare science: how anthropogenic landscape transformations affect not just population-level metrics, but also the welfare of individual wild animals. It provides a methodological proof of concept by evaluating a multi-domain suite of non-invasive behavioral, physiological, and health-related indicators that can be validated under real-world field conditions, explicitly assessing their sensitivity, directionality, and cross-domain convergence across contrasting anthropogenic contexts.
Assessing anthropogenic fear in a wild decapod crustacean
Grantee: Mark Briffa
Institution: University of Plymouth, United Kingdom
Project summary
This project will investigate whether urban coastal environments alter fear-related behavioral and physiological responses in wild hermit crabs, using two species that differ in their history of exposure to urbanization. Researchers will compare crabs of both species collected from urban and rural shorelines and measure indicators that may be linked to affective state, including startle responses, shell investigation behavior, and resting metabolic rate under field conditions. By repeatedly measuring behavior within individuals, they will also test whether urban disturbance influences behavioral predictability, which may provide additional insight into fear and stress-related processes.
Grantee: Mark Briffa
Institution: University of Plymouth, United Kingdom
Grant amount: $49,656
Grant type: Discovery grants
Focal species: Common European hermit crab (Pagurus bernhardus), St. Pirin’s hermit crab (Clibanarius erythropus)
Conservation status: Not evaluated
Disciplines: Animal behavior, climate science
Research location: United Kingdom
Project summary
This project will investigate whether urban coastal environments alter fear-related behavioral and physiological responses in wild hermit crabs, using two species that differ in their history of exposure to urbanization. Researchers will compare crabs of both species collected from urban and rural shorelines and measure indicators that may be linked to affective state, including startle responses, shell investigation behavior, and resting metabolic rate under field conditions. By repeatedly measuring behavior within individuals, they will also test whether urban disturbance influences behavioral predictability, which may provide additional insight into fear and stress-related processes.
Why we funded this project
This project applies concepts and analytical approaches from animal personality research to the assessment of welfare-related states in free-living animals. It also addresses a priority taxonomic gap in the field: Crustaceans are abundant yet remain largely neglected in wild animal welfare research, despite growing evidence for their behavioral complexity and sentience-related capacities.
Validating behavioral, acoustic, and physiological indicators of welfare in urban birds
Grantee: Miriam Soledad Vazquez
Institution: Universidad Nacional del Sur, Argentina
Project summary
This project will evaluate whether behavioral, acoustic, and physiological indicators consistently reflect welfare-relevant experiences in two urban thrush species. Welfare will be assessed by combining focal behavioral observations (vigilance, aggression, displacement, foraging activity, and tolerance of conspecifics and heterospecifics), passive acoustic monitoring (focal activity rates and calling patterns), and feather glucocorticoid analyses across urban sites that differ in vegetation cover and human disturbance. These indicators will be interpreted together to evaluate whether they show concordant, biologically meaningful responses across urban environments, while recognizing that different indicators may capture different temporal or contextual dimensions of affective state and welfare.
Grantee: Miriam Soledad Vazquez
Institution: Universidad Nacional del Sur, Argentina
Grant amount: $9,600
Grant type: Seed grants
Focal species: Patagonian thrush (Turdus falcklandii), Rufous-bellied thrush (T. rufiventris)
Conservation status: N/A: Multiple focal species
Disciplines: Ornithology, animal behavior, physiology
Research location: Argentina
Project summary
This project will evaluate whether behavioral, acoustic, and physiological indicators consistently reflect welfare-relevant experiences in two urban thrush species that have recently expanded their ranges into Argentine cities. Welfare will be assessed by combining focal behavioral observations (vigilance, aggression, displacement, foraging activity, and tolerance of conspecifics and heterospecifics), passive acoustic monitoring (focal activity rates and calling patterns), and feather glucocorticoid analyses across urban sites that differ in vegetation cover and human disturbance. These indicators will be interpreted together to evaluate whether they show concordant, biologically meaningful responses across urban environments, while recognizing that different indicators may capture different temporal or contextual dimensions of affective state and welfare.
Why we funded this project
The study provides a proof of concept for integrating behavioral observations, passive acoustic monitoring, and physiological measures within a multidimensional welfare assessment framework applied under natural conditions. It also addresses whether commonly proposed welfare indicators remain informative across different species and environmental contexts, including urban environments shaped by human disturbance and novel biotic interactions.